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Future Research Directions and Unanswered Questions Despite 50+ years since Pickarts initial isolation work, significant mechanistic and clinical questions remain: Receptor identity: GHK-Cus gene regulatory effects are well-documented, but the specific cellular receptors mediating them remain incompletely characterized

At the mechanistic level, H exerts neuroprotective effects through three core mechanisms: (1) selective scavenging of toxic ROS, such as -OH and ONOO-, along with the activation of the Nrf2/ARE pathway to enhance endogenous antioxidant defenses

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