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GSH can also serve as a substrate for S-glutathionylation of proteins in the presence of small redox enzyme glutaredoxins (GRX), which also utilizes GSH as a cofactor ( Figure 1 )

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Conversely, under hypoxic conditions, phosphoglycerate mutase 5 (PGAM5) and ULK1 catalyze the dephosphorylation and phosphorylation of Ser13 and Ser17, respectively, thereby inducing mitophagy [19]

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